Unterfamilie Oxytelinae Coleoptera - Staphylinidae
  By Gy. Makranczy (Rearranged by Arved Lompe)
Informieren Sie mich bitte über Fehler oder Ergänzungen über mailbox@lompe.de 		Makranczy, Gy. (2006): Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae). - Annales historico-naturales musei Nationalis Hungarici (Budapest), 98:29-120 Private Datei: F:\taxa\coleo\scans\Makranczy2006.pdf
  This key is not intended to be a practical identification guide, rather a systematic one, which helps workers determine whether a particular species belongs to a given genus or not. This goal can only be achieved by often relying on anatomical characters (necessitating permanent preparations or dissections). Since the superficial similarity between unrelated taxa can be great, it is unavoidable that less experienced workers examine at least a few of the key features of the genital segments, which can unequivocally determine the placement of a specimen. A more practical approach would require the use of external morphological characters that can vary and result in confusion - not counting the (great) possibility of previously unseen species possessing yet undiscovered character states.  
#1 Abdominal segments III to VI with one pair of laterosclerites per segment. (Deleasterini)

   ...2

  
-- Abdominal segments III to VI with two pairs of laterosclerites per segment

   ...7

  
#2 Procoxal fissure fully open. Tibia with longitudinal row of spines.

   ...3

  
-- Procoxal fissure nearly or quite closed. Tibia without longitudinal row of spines absent

   ...4

  
#3 Pronotum distinctly convex in cross-section, laterally not margined; elytra without rows of punctures. Tergite VIII with serrate posterior margin. Body surface more or less dull. Habitus [Abb.1].

   ...Deleaster Er.

DELEASTER DICHROUS
Abb.1		 
-- Pronotum flat and distinctly margined laterally; elytra with rows of punctures. Tergite VIII with smooth posterior margin. Body surface shining, except for the impressions and rows of punctures on head, pronotum, and elytra [Abb.2]

   ...Platydeleaster
PLATYDELEASTER BIMACULATUS
Abb.2		 
#4 Body, especially dorsum, with numerous scale-like setae

   ...5

  
-- Body without scale-like setae

   ...6

  
#5 Scale-like setae not plumose. Tarsal formula 5-5-5

   ...Oxypius

  
-- Scale-like setae plumose. Tarsal formula 3-3-3

   ...Euphanias

  
#6 Labrum with anterior margin emarginate. Elytra (from apex of scutellum) at least as long as pronotum. Habitus [Abb.3].

   ...Syntomium Curt.

SYNTOMIUM AENEUM
Abb.3		 
-- Labrum with anterior margin broadly convex; elytra (from apex of scutellum) distinctly shorter than pronotum [Abb.4]

   ...Mitosynum
MITOSYNUM VOCKEROTHI
Abb.4		 
#7 Reduced second sternite with a very short, immobile membranous connection to third sternite (hence only 6 complete sterna can be counted) (Coprophilini)

   ...8

  
-- Well developed second sternite articulated to third by a long connecting membrane (hence 7 complete sterna can be counted)

   ...10

  
#8 Gular sulci separated along entire length [Abb.5]

   ...Coprophilus Latr.

TEROPALPUS LUNIGER
Abb.5		 
-- Gular sulci confluent anteriorly

   ...9

  
#9 Epistomal sulcus present [Abb.6]

   ...Coprostygnus
COPROSTYGNUS OPTANDUS
Abb.6		 
-- Epistomal sulcus absent

   ...Homalotrichus

  
#10 Eyes extending onto ventral portion of head [Abb.7] (Planeustomini)

   ...11
MANDA MANDIBULARIS
Abb.7		 
-- Eyes not extending onto ventral portion of head

   ...12

  
#11 Five distinct tarsomeres. Brick-wall pattern in intersegmental membrane. Habitus [Abb.8].

   ...Manda Blackw. 1952

MANDA MANDIBULARIS
Abb.8		 
-- Four tarsomeres with additional non-setose pseudoarticle [Abb.9], brickwall pattern in intersegmental membrane absent. Habitus [Abb.10].

   ...Planeustomus duVal

PLANEUSTOMUS PALPALIS
Abb.9		PLANEUSTOMUS PALPALIS
Abb.10
#12 Antennae geniculate (flexing posteriorly) (Blediini)

   ...13

  
-- Antennae not geniculate (Oxytelini)

   ...14

  
#13 Setation behind the front of labrum hyperdeveloped so that the lateral setae reach well beyond four times the length of labrum. Habitus [Abb.11]

   ...Eppelsheimius
EPPELSHEIMIUS PIRAZZOLII
Abb.11		 
-- Setation behind the front of labrum is well developed but not reach beyond four times the length of labrum. Habitus [Abb.12].

   ...Bledius Leach in Samouelle, 1819

BLEDIUS GALLICUS
Abb.12		 
#14 Tarsomeres five

   ...15

  
-- Tarsomeres less than five

   ...25

  
#15 Elytra long, almost entirely covering abdomen

   ...Sartallus

  
-- Elytra short, abdomen almost fully visible

   ...16

  
#16 Pronotum cylindrical, lateral margins invisible from dorsal view [Abb.13]

   ...Mimopaederus
MIMOPAEDERUS INSULARIS
Abb.13		 
-- Pronotum with lateral margins always visible from dorsal view

   ...17

  
#17 Gular sulci narrowly separated anteriorly, more widely posteriorly [Abb.5]

   ...Teropalpus
TEROPALPUS LUNIGER
Abb.5		 
-- Gular sulci fused at least partially [Abb.14]

   ...18
BLEDIOTROGUS GUTTIGER
Abb.14		 
#18 Gular sulci not fused anteriorly, but merged posteriorly [Abb.14]

   ...Blediotrogus
BLEDIOTROGUS GUTTIGER
Abb.14		 
-- Gular sulci always fused anteriorly

   ...19

  
#19 Labrum undivided

   ...20

  
-- Labrum divided along mid-line [Abb.15]

   ...21
THINODROMUS JUANFERNANDEZIANUS
Abb.15		 
#20 Basolateral portions of tergite X fused to tergites IX (rhomboid fusion) [Abb.16]. Habitus [Abb.17]

   ...Xerophygus
XEROPHYGUS HREBLAYI
Abb.16		XEROPHYGUS HREBLAYI
Abb.17
-- Tergite X always complete, unmodified

   ...Ochthephilus Mulsant & Rey 1856

 
#21 Antennae not extremely elongated (mid-antennal segments less than three times as long as broad). Habitus [Abb.18] [Abb.19]

   ...Thinodromus Kraatz 1857

THINODROMUS SCHILLHAMMERI
Abb.18		THINODROMUS JUANFERNANDEZIANUS
Abb.19
-- Antennae extremely elongated (mid-antennal segments more than three times as long as broad) [Abb.20]

   ...Trogactus
TROGACTUS OCELLATUS
Abb.20		 
#22 Tarsomeres four

   ...23

  
-- Tarsomeres less than four

   ...24

  
#23 Basal segment of labial palp with strong setation mediad. Distal segment of maxillary palp much longer than penultimate. Habitus [Abb.21]

   ...Crassodemus
CRASSODEMUS FORAMINOSUS
Abb.21		 
-- Basal segment of labial palp without such setation. Distal segment of maxillar palp quite reduced, much smaller and thinner than penultimate

   ...Pareiobledius

  
#24 Tarsomeres three.

   ...25

  
-- Tarsomeres two.

   ...34

  
#25 Basal two tarsomeres strongly compressed. Habitus [Abb.22]

   ...Carpelimus Leach in Samouelle, 1819

CARPELIMUS BIFIDUS
Abb.22		 
-- Basal two tarsomeres not compressed

   ...26

  
#26 Labrum medially divided [Abb.15]

   ...27
THINODROMUS JUANFERNANDEZIANUS
Abb.15		 
-- Labrum not divided medially

   ...28

  
#27 Tergum X not bilobed. Tergal basolateral ridges absent, epistomal sulcus present, sometimes accentuated by dark line

   ...Aploderus Steph.

 
-- Tergum X slightly bilobed at apex. Tergal basolateral ridges present, epistomal sulcus absent

   ...Paraploderus

  
#28 Labrum anteriorly with a very conspicuous rounded incision

   ...Parosus

  
-- Labrum without such incision

   ...29

  
#29 Pronotal lateral marginal bead absent

   ...Ecitoclimax

  
-- Pronotal marginal bead present.

   ...30

  
#30 Antennomeres nine

   ...Jerozenia

  
-- Antennomeres eleven

   ...31

  
#31 Elytra slightly overlapping at suture

   ...Platystethus Mannh.

 
-- Elytra strictly parallel at suture

   ...32

  
#32 Scutellar impression distinctly diamond-shaped. Aedeagus with apicomedial hook

   ...Oxytelus Gravenhorst

 
-- Scutellar impression different. Aedeagus without apicomedial hook

   ...33

  
#33 Neck about half as wide as head at temples

   ...Apocellus

  
-- Neck wider than half of head width at temples [Abb.23]

   ...Anotylus Thomson 1859

ANOTYLUS SETICORNIS
Abb.23		 
#34 Elytral epipleural ridge present [Abb.24]. Hypomeron of pronotum well developed, concealing most of protrochantin [Abb.25]

   ...Neoxus
NEOXUS CRASSICORNIS
Abb.24		NEOXUS CRASSICORNIS
Abb.25
-- Elytral epipleural ridge absent [Abb.26]. Hypomeron of pronotum less extended mediad; protrochantin visible

   ...35
THINOBIUS APFELBECKI
Abb.26		 
#35 Scutellum pubescent (to various degrees)

   ...Thinobius Kiesw.

 
-- Scutellum without pubescence [Abb.27]

   ...Sciotrogus
SCIOTROGUS OPACUS
Abb.27		 
  Anotylus
Aploderus
Apocellus
Blediotrogus
Bledius
Carpelimus
Coprophilus
Coprostygnus
Crassodemus
Deleaster
Ecitoclimax
Eppelsheimius
Euphanias
Homalotrichus
Jerozenia
Manda
Mimopaederus
Mitosynum
 Neoxus
Ochthephilus
Oxypius
Oxytelus
Paraploderus
Pareiobledius
Parosus
Planeustomus
Platydeleaster
Platystethus
Sartallus
Sciotrogus
Syntomium
Teropalpus
Thinobius
Thinodromus
Trogactus
Xerophygus
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