Helophorus_mh Bestimmungstabelle

Genus Helophorus F., 1775 after M. Hansen

Coleoptera

By Michael Hansen (Rearranged by Arved Lompe)

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Key to the species of Fennoscandia and Denmark.

Intercalary stria present.

...2

Intercalary stria absent.

...10

Elytra entirely black, with raised tubercles on 3rd, 5th and 7th interstice. Aedoeagus [fig.1].

...tuberculatus Gyll., 1808

fig.1

Elytra brown or yellowish, without tubercles.

...3

Alternate interstices of elytra markedly raised above the others. Pseudepipleura opposite metacoxae as wide as, or wider than epipleura [fig.2] [fig.3].

...4

fig.2

fig.3

Alternate interstices of elytra at most weakly raised above the others. Pseudepipleura opposite metacoxae narrower than epipleura.

...7

Terminal segment of maxillary palpi asymmetrical, with straighter inner face [fig.4]. Alternate interstices of elytra more moderately ridged. Pseudepipleura opposite metacoxae about as wide as epipleura [fig.2]. Aedoeagus [fig.5].

...sibiricus Motschulsky

fig.4

fig.2

fig.5

Terminal segment of maxillary palpi symmetrical [fig.6]. Alternate interstices of elytra strongly ridged. Pseudepipleura opposite metacoxae about twice as wide as epipleura [fig.3].

...5

fig.6

fig.3

Smaller, 3.0-4.0 mm. Elytra at least 1/2 longer than wide. Internal intervals of pronotum rather weakly narrowed behind middle [fig.7]. Aedoeagus [fig.8].

...nubilus F., 1777

fig.7

fig.8

Larger, 4.0-5.6 mm. Elytra only about 2/5 longer than wide. Internal intervals of pronotum strongly narrowed behind middle ( [fig.9] [fig.10]).

...6

fig.9

fig.10

Sides of pronotum distinctly sinuate posteriorly. Elytra with sharp humeral angle [fig.10]. Aedoeagus [fig.11].

...rufipes (Bosc., 1791)

fig.10

fig.11

Sides of pronotum evenly and rather feebly curved, not sinuate posteriorly. Humeral angle rounded [fig.9]. Aedoeagus [fig.12].

...porculus Bedel, 1881

fig.9

fig.12

Head and pronotum strongly and evenly granulate, the granules very distinct and strikingly well defined, show ing no trace of fusing. Posterior margin of last visible abdominal sternite fairly strongly dentate [fig.13]. Aedoeagus [fig.14].

...grandis Ill., 1798

fig.13

fig.14

Head and pronotum more feebly granulate, the granules to some extent fusing and indistinct, especially on internal intervals of pronotum. Posterior margin of last visible abdominal sternite with finer (or indistinct) dention [fig.15].

...8

fig.15

Frontal furrow narrowly linear (or sublinear), rather deep. Elytra about 1/2 longer than wide. Sides of pronotum normally markedly sinuate posteriorly. Aedoeagus [fig.16].

...strandi Angus

fig.16

Frontal furrow almost always expanding anteriorly. Elytra narrower, normally about 3/5 longer than wide. Sides of pronotum normally more weakly sinuate.

...9

Elytra dark brown, often with distinct lighter mottling. ♂: Parameres (dorsal view) making up less than half the total aedeagal length (index parameres: aedeagus = 0.42-0.47), their outer face normally distinctly rounded [fig.17]. ♀: 9th horseshoe-shaped tergite (retracted in abdomen) with distinct lateral projections basal to which sides are distinctly concave [fig.18].

...aquaticus (L., 1758)

fig.17

fig.18

Elytra often slightly paler, usually without distinct lighter mottling. ♂: Parameres (dorsal view) making up about half of the total aedeagal length (index parameres: aedeagus = 0.47-0.51), their outer face straighter, often feebly sinuate [fig.19]. ♀: 9th tergite on the average a little narrower, with less distinct lateral projections, often almost straight-sided [fig.20].

...aequalis Thoms., 1868

fig.19

fig.20

#10

Legs and palpi very dark, often black, with distinct metallic reflections. Aedoeagus [fig.21].

...glacialis Villa, 1833

fig.21

Legs and palpi reddish or yellowish (sometimes brown), without any trace of metallic reflections.

...11

#11

Terminal segment of maxillary palpi symmetrical oval [fig.22] [fig.23].

...12

fig.22

fig.23

Terminal segment of maxillary palpi asymmetrical, with straighter inner face [fig.24].

...13

fig.24

#12

Pronotum moderately convex, lateral margins straight (or only weakly sinuate)in posterior half [fig.25]. Aedoeagus [fig.26].

...brevipalpis Bedel, 1881

fig.25

fig.26

Pronotum strongly convex, lateral margins markedly sinuate posteriorly [fig.27]. Aedoeagus [fig.28].

...arvernicus Muls., 1846

fig.27

fig.28

#13

Antennae 8-segmented [fig.29]. Frontal furrow narrowly linear (except in some rare specimens of redtenbacheri).

...14

fig.29

Antennae 9-segmented [fig.30]. Frontal furrow narrowly linear or often wider, often expanded anteriorly..

...16

fig.30

#14

Larger, 4.5-5.0 mm. Ground colour reddish brown or brown. Aedoeagus [fig.31].

...pallidus Gebler

fig.31

Smaller, 2.3-3.0 mm. Ground colour black, elytra paler.

...15

#15

Pronotum [fig.32] smooth and shining, often almost polished, normally finely punctured (almost without granulation). Head with a distinct pair of accessory furrows running (almost) parallel to median frontal furrow, between it and eyes [fig.33]. Aedoeagus [fig.34].

...nanus Sturm, 1836

fig.32

fig.33

fig.34

Pronotum rather coarsely granulate (though granules rather low). Head without such accessory furrows. Aedoeagus [fig.35].

...redtenbacheri Kuw., 1885

fig.35

#16

Elytra with distinct (often very conspicuous) paler spots. Pseudepipleura broadly visible from below, opposite metacoxae about equally wide as epipleura (as [fig.2]). Frontal furrow broadly linear [fig.36] or expanded anteriorly (as [fig.37]).

...17

fig.2

fig.36

fig.37

Elytra normally without distinct (or at most vaguely defined) paler spots. If pseudepipleura are broadly visible from below, frontal furrow is narrowly linear [fig.38].

...18

fig.38

#17

Elytra about 1/2 longer than wide, very strongly striate, interstices strongly convex, in part obtusely ridged; without large conspicuous pale spot behind arrow-mark. Frontal furrow not expanded anteriorly. Aedoeagus [fig.39].

...croaticus Kuw., 1886

fig.39

Elytra 3/5 longer than wide, rather strongly striate, interstices much less convex, not ridged. Lighter mottling of elytra more extensive, normaly forming a large conspicuous pale spot on each elytron, behind arrow-mark. Frontal furrow expanded anteriorly. Aedoeagus [fig.40].

...dorsalis (Marsh., 1802)

fig.40

#18

Head and pronotum strongly and evenly granulate, the granules very distinct and strikingly well defined, also on internal intervals of pronotum, showing no trace of fusing. Frontal furrow normally rather narrow, sublinear [fig.41]. Rather large, dark species, normally with a faint greenish hue on elytra; length 3.8-4.8 mm. Aedoeagus [fig.42].

...asperatus Rey, 1885

fig.41

fig.42

Granulation on middle portion of pronotum almost always somewhat reduced, only seldom with distinct granules on internal intervals (the granules then fairly low and not strikingly well defined, except in some very rare specimens of obscurus, cf. the description of that species). Frontal furrow normally either wider and expanded anteriorly [fig.37] or narrowly linear [fig.38].

...19

fig.37

fig.38

#19

Elytra with distinct, though sometimes faint greenish, purplish or bronze hue, almost always dark. Elytra with a distinct oblique impression in anterior third.

...20

Elytra without any trace of metallic reflections, and without distinct impression in anterior third.

...21

#20

Submedian furrow of pronotum rather weakly curved outwards in middle, not distinctly angular [fig.43]. Elytra normally tapered at apex. ♂: Aedeagus [fig.44].

...flavipes F., 1792

fig.43

fig.44

Submedian furrow of pronotum normally more strongly and markedly angularly curved outwards in middle [fig.45]. Elytra on the average a little wider, apex usually more bluntly rounded. ♂: Aedeagus [fig.46].

...obscurus Muls., 1844

fig.45

fig.46

#21

Frontal furrow narrowly linear, rather deep [fig.38]. Pronotum strongly convex, also distinctly convex longitudinally, or internal intervals strongly convex about middle, somewhat humpy .

...22

fig.38

Frontal furrow appearing more shallow, normally expanded anteriorly (as [fig.37]). Pronotum less strongly convex, hardly convex longitudinally, internal intervals not humpy.

...24

fig.37

#22

Pseudepipleura narrow, not visible from below (as [fig.47]). Pronotum rather evenly convex, with uniform, close and distinct, but rather feeble granulation; granules normally visible also on internal intervals (though often slightly weaker). Length, 3.0-4.0 mm. Aedoeagus [fig.48].

...laticollis Thoms., 1853

fig.47

fig.48

Pseudepipleura broadly visible from below, opposite metacoxae aboùt as wide as epipleura (as [fig.2]). Internal intervals of pronotum more or less humpy, normally with strongly reduced granulation, or even simply punctured.

...23

fig.2

#23

Larger, 3.5-4.2 mm. Pronotum [fig.49] widest just be fore middle, the sides a little straightened in posterior half; strongly convex (though longitudinally only slightly so), internal intervals usually markedly humpy. Aedoeagus [fig.50].

...strigifrons Thoms., 1868

fig.49

fig.50

Smaller, 2.5-3.2 mm. Pronotum widest just behind middie, the sides evenly rounded (almost as in [fig.32]); strongly arched and distinctly convex longitudinally. Aedoeagus [fig.51].

...pumilio Er., 1837

fig.32

fig.51

#24

Elytra yellowish brown, rather strongly striate. Pronotum [fig.52] strongly, rather evenly granulate; granules on internal intervals rather feeble, but usually distinct. Metatarsal claw segment (excluding claws) distinctly longer than 1st + 2nd segment [fig.53]. Length 3.2-4.5 mm. Aedoeagus [fig.54].

...fulgidicollis Motsch., 1860

fig.52

fig.53

fig.54

Elytra paler, striae finer (except granularis: 2.2-2.8 mm). Pronotum usually more weakly granulate, especially on internal intervals (except granularis and some lapponicus). Metatarsal claw segment (excluding claws) not longer than 1st + 2nd segment [fig.55] (except lapponicus).

...25

fig.55

#25

Granulation of pronotum very reduced, at most visible on external intervals (and only weakly so). Pronotum very shining, bright metallic. Elytra very pale, finely striate. ♂: Aedeagus characteristic [fig.56].

...longitarsis Woll., 1864

fig.56

Granulation of pronotum more distinct, also distinct on middle intervals.

...26

#26

Pronotum rather strongly narrowed posteriorly [fig.57], the sides straightened posteriorly and normally feebly sinuate; pronotal granulation rather fine, on internal intervals usually reduced to simple punctuation. Elytra pale, finely striate. ♂: Aedeagus characteristic [fig.58].

...griseus Hbst., 1793

fig.57

fig.58

Pronotum a little less narrowed posteriorly, the sides more evenly rounded, usually also posteriorly (though here often a little straighter).

...27

#27

Metatarsal claw segment (excluding claws) distinctly longer than 1st + 2nd segment (as [fig.53]). Size on the average larger, 3.3-4.0 mm. ♂: Aedeagus [fig.59].

...lapponicus Thomson

fig.53

fig.59

Metatarsal claw segment (excluding claws) not longer than 1st + 2nd segment [fig.55]. Size on average smaller, 2.2-3.6 mm.

...28

fig.55

#28

Pronotum [fig.60] rather strongly convex, with rather even, close and usually coarse, though very low granulation. Marginal furrows narrow, hardly paler. Submedian furrow only rarely angled in middle. Small, 2.2-2.8 mm. ♂: Aedeagus [fig.61].

...granularis (L., 1761)

fig.60

fig.61

Pronotum [fig.62] normally weaker convex, granulation (especially on internal intervals) weaker. Marginal furrow wider and often paler; if marginal furrow is not istinctly paler, submedian furrow is usually distinctly angled in middle. On average larger, 2.8-3.6 mm .

...29

fig.62

#29

Pronotum a little more convex and rather strongly granulate, the margins reddish or brownish, not conspicuously paler. ♂: Aedeagus [fig.63].

...discrepans Rey

fig.63

Pronotum [fig.62] a little less convex, normally more weakly granulate, anterior and lateral margins normally distinctly paler, usually broadly yellow. ♂: Aedeagus [fig.64]. Habitus [fig.65].

...minutus F., 1775

fig.62

fig.64

fig.65

aequalis
aquaticus
arvernicus
asperatus
brevipalpis
croaticus
discrepans
dorsalis
flavipes
fulgidicollis
glacialis
grandis
granularis
griseus
lapponicus

laticollis
longitarsis
minutus
nanus
nubilus
obscurus
pallidus
porculus
pumilio
redtenbacheri
rufipes
sibiricus
strandi
strigifrons
tuberculatus

Erstellt am: 15.07.2024
Letzte Aktualisierung: 21.07.2024 - 13:48:30
Mit Determin (V4.2.298-4) von: Arved Lompe
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