Genus Miarus - Europa Coleoptera - Rhynchophora - Curculionidae - Mecininae
  By Roberto Caldara (sligthly modified by Arved Lompe) If you find any errors or omissions please mail me at mailbox@lompe.de
  Caldara, R. (2001): Phylogenetic analysis and higher classification of the tribe Mecinini (Coleoptera: Curculionidae, Curculioninae). Kol. Rundschau 71:171-203.
Caldara, R. (2007): Taxonomy and phylogeny of the species of the weevil genus Miarus Schönherr, 1826. (Coleoptera: Curculionidae, Curculioninae). - Koleopterologische Rundschau, 77:199-248 Private Datei: F:\taxa\coleo\scans\Caldara2007 Miarus.tif
 
#1 ♂: Pygidium simple, without median gibbosity and fovea; ventrite 5 without fovea and teeth; elytra somewhat long [fig.1]; body length 1.9-2.5 mm.
♀: Pygidium simple and bursa copulatrix without sclerites.

   ...2

MIARUS AFER
fig.1
 
-- ♂: Pygidium with more or less pronounced median gibbosity and fovea; ventrite 5 without or with more or less deep fovea and two more or less robust teeth laterally to fovea [fig.2]; elytra fairly short to short [fig.3]; body length 1.6-3.5 mm.
♀: Pygidium with fovea and bursa copulatrix with two semilunate sclerites.

   ...3

MIARUS CAMPANULAE
fig.2
MIARUS MONTICOLA
fig.3
#2 Basal margin of elytra nearly regularly and weakly concave, scales of elytral interstriae nearly completely arranged in single regular rows; in ♂ uncus of protibia small [fig.4]; median lobe of aedeagus elongate and narrow, with short apex, endophallus with elongate flagellum and without elongate median ventral sclerite [fig.5]. Morocco, Algeria, Tunisia.

   ...afer Daniel, 1912

MIARUS AFER
fig.4
MIARUS AFER.GIF
fig.5
-- Basalrand der Flügeldecken von der Naht bis zum 5. Streifen leicht konvex dann deutlich schräg nach vorne verlaufend. Schuppen der Flügeldeckenstreifen 1-2 reihig angeordnet. ♂: Voderschienen mit großem Spitzendorn. Medianlobus des Aedoeagus kürzer und breiter mit langer Spitze; Endophallus mit kurzem Flagellum und langem, ventralem Sklerit.
Basal margin of elytra weakly convex from suture to fifth interstria, then distinctly directed forwards; scales of elytral interstriae arranged in 1-2 irregular rows; in ♂ uncus of protibia robust [fig.6]; median lobe of aedeagus shorter and wide, with long apex, endophallus with short flagellum and with elongate median ventral sclerite [fig.7]. Greece, Turkey, Syria, Jordan.

   ...praecursor Caldara, 2001

MIARUS PRAECURSOR
fig.6
MIARUS PRAECURSOR.GIF
fig.7
#3 Ventrite 5 of ♂ at most with weak depression, covered with scarce to dense hair-like vestiture (Figs. 34-36).

   ...4

 
-- Ventrite 5 of ♂ with fovea moderately shallow to deep, nearly glabrous (Figs. 37-40).

   ...6

 
#4 Body length 1.8-2.3 mm; protibiae short, in male with very robust uncus as long as 1/5 of tibia [fig.8]; depression of ventrite 5 of male with scarce vestiture [fig.9]; median lobe of aedeagus in dorsal view distinctly wider at base than at apex, in lateral view as thin at base as at apex, with orifice very small and narrow [fig.10]; habitus as in [fig.11]. Croatia, Macedonia, Greece, Turkey, Armenia, Syria.

   ...stoeckleini Franz, 1947

MIARUS STOECKLEINI
fig.8
MIARUS STOECKLEINI
fig.9
MIARUS STOECKLEINI.GIF
fig.10
MIARUS STOECKLEINI
fig.11
-- Body length 2.0-2.8 mm; protibiae longer, in male with moderately robust uncus as long as 1/8 of tibia; depression of ventrite 5 of male with vestiture denser than at sides [fig.12]; median lobe of aedeagus in dorsal view as wide at base as at apex, in lateral view distinctly robust at apex, with orifice larger [fig.13].

   ...5

MIARUS ROTUNDICOLLIS
fig.12
MIARUS ROTUNDICOLLIS.GIF
fig.13
#5 Pronotum 1/3 wider than long, usually widest near middle; basal margin of elytra regularly concave [fig.14]; ventrite 5 of male without posterolateral teeth; median lobe of aedeagus without distinct ventral tongue [fig.13], endophallus [fig.15]. Albania, Macedonia, Greece.

   ...rotundicollis Desbrochers, 1893

MIARUS ROTUNDICOLLIS
fig.14
MIARUS ROTUNDICOLLIS
fig.15
-- Pronotum 1/2 wider than long, widest at basal 1/3; basal margin of elytra usually weakly convex to fifth interstria then distinctly directed forwards [fig.16]; ventrite 5 of male usually with two small posterolateral teeth; median lobe of aedeagus with distinct ventral tongue [fig.17]. Bulgaria, Greece.

   ...hellenicus Dieckmann, 1978

MIARUS HELLENICUS
fig.16
MIARUS HELLENICUS.GIF
fig.17
#6 Ventrite 5 of male with fovea moderately shallow and teeth usually moderately robust [fig.18].

   ...7

MIARUS ABNORMIS
fig.18
 
-- Ventrite 5 of male with fovea deep and teeth usually very robust [fig.19].

   ...11

MIARUS URSINUS
fig.19
 
#7 Ventrite 5 of male with a very small shining median tooth anterior to fovea [fig.20].

   ...8

MIARUS ARAXIS
fig.20
 
-- Ventrite 5 of male without tooth anterior to fovea.

   ...9

 
#8 Scales of dorsal vestiture brown, those covering elytral interstriae recumbent [fig.21]; rostrum in lateral view distinctly curved, in female only slightly longer than in male; ventrite 5 of male with shallower fovea; median lobe of aedeagus with tip short, not visible in dorsal view [fig.22]. Armenia, Georgia, Turkey.

   ...araxis Reitter, 1907

MIARUS ARAXIS
fig.21
MIARUS ARAXIS.GIF
fig.22
-- Scales of dorsal vestiture white, those covering elytral interstriae suberect to erect [fig.23]; rostrum in lateral view in male weakly curved, in female nearly straight [fig.24] and somewhat longer than in male; fovea of ventrite 5 deeper; median lobe of aedeagus with tip long, clearly visible in dorsal view [fig.25]. Armenia, Iran.

   ...dentiventris Reitter, 1907

MIARUS DENTIVENTRIS
fig.23
MIARUS DENTIVENTRIS
fig.24
MIARUS DENTIVENTRIS.GIF
fig.25
 
#9 Rostrum in lateral view distinctly tapered in apical 1/3 [fig.26]; body length 2.1-2.6 mm; scales of dorsal vestiture whitish [fig.27]. Ukraine, Turkey.

   ...solarii Smreczynski, 1957

MIARUS SOLARII
fig.26
MIARUS SOLARII
fig.27
-- Rostrum in lateral view nearly of same width from base to apex; body length 2.4-3.0; scales of dorsal vestiture whitish to brown.

   ...10

 
#10 Scales of dorsal vestiture whitish [fig.28]; median lobe of aedeagus without flanges at apex laterally [fig.29]. Finland, north-western Russia (Karelia, Arkhangel'sk Prov., Leningrad Prov., Kirovsk Prov., Bryansk, Moscow Prov.; ZISP), Ukraine, Poland, Germany, Czech Republic, Slovakia, Austria, Bosnia and Herzegovina, Serbia, Albania, Montenegro, Bulgaria, Greece

Meldungen in Google Earth anzeigen   ...monticola Petri, 1912

MIARUS MONTICOLA
fig.28
MIARUS MONTICOLA3.GIF
fig.29
-- Scales of dorsal vestiture whitish to brown or of both colours intermixed; median lobe of aedeagus with distinct flanges at apex laterally [fig.30]. South-eastern Poland, Austria, north-eastern Italy, Slovenia, Croatia, Serbia, Montenegro, Macedonia. Its presence in southern France needs confirmation.

   ...abnormis Solari, 1947

MIARUS ABNORMIS2.GIF
fig.30
 
#11 Rostrum in female very long (Rl/Pl > 1.30) and distinctly longer than in male [fig.31] [fig.32] [fig.33].

   ...12

MIARUS BANATICUS
fig.31
MIARUS LONGICOLLIS
fig.32
MIARUS ABEILLEI
fig.33
 
-- Rostrum in female only slightly longer than in male (Rl/Pl < 1.05).

   ...14

 
#12 Scales of elytral interstriae arranged in 1-2 irregular rows, subrecumbent; median lobe of aedeagus at apex with small auricles (au) and very small mitre (m) [fig.34]; habitus of female as in [fig.35]. Athos, Greece. Only known from the type locality.

   ...Iongicollis

MIARUS LONGICOLLIS.GIF
fig.34
MIARUS LONGICOLLIS
fig.35
-- Scales of elytral interstriae mainly arranged in 3-4 irregular rows, recumbent; median lobe of aedeagus at apex with distinct auricles and mitre.

   ...13

 
#13 Rostrum nearly straight in lateral view in both sexes [fig.33] and less punctate in male; scales of dorsal vestiture whitish to brown; median lobe of aedeagus with auricles distinctly uncinate and without spines [fig.36]; habitus of female as in [fig.37]. South-eastern France, Switzerland, north-western Italy.

   ...abeillei Desbrochers, 1893

MIARUS ABEILLEI
fig.33
MIARUS ABEILLEI.GIF
fig.36
MIARUS ABEILLEI
fig.37
 
-- Rostrum moderately curved in lateral view in both sexes [fig.31] and more punctate in male; scales of dorsal vestiture usually brown or white and brown intermixed; median lobe of aedeagus with auricles not uncinate but usually with some minute spines [fig.38]; habitus of female as in [fig.39]. Bosnia and Herzegovina, Romania.

   ...banaticus Reitter, 1907

MIARUS BANATICUS
fig.31
MIARUS BANATICUS.GIF
fig.38
MIARUS BANATICUS
fig.39
 
#14 Median lobe of aedeagus with medium sized [fig.40] to large auricles [fig.41].

   ...15

MIARUS URSINUS.GIF
fig.40
MIARUS CAMPANULAE3
fig.41
-- Median lobe of aedeagus at most with small auricles [fig.42] [fig.43].

   ...17

MIARUS SIMPLEX.GIF
fig.42
MIARUS AJUGAE3
fig.43
#15 Ventrite 5 of male with fovea usually very deep and teeth distinctly robust [fig.2]; median lobe of aedeagus with auricles large, distinctly uncinate [fig.41]; body medium to large sized (length 2.3-3.4 mm); scales of elytral interstriae arranged in more than one irregular rows [fig.44]; uncus of protibia in male moderately robust [fig.45]. Great Britain, Ireland, Spain, France, Belgium, Holland, Poland, Germany, Hungary, Czech Republic, Slovakia, Austria, peninsular Italy, Denmark, Norway, Sweden, Finland, western Russia.

Meldungen in Google Earth anzeigen   ...campanulae (L., 1767)

MIARUS CAMPANULAE
fig.2
MIARUS CAMPANULAE
fig.44
MIARUS CAMPANULAE
fig.45
 
-- Ventrite 5 of male with fovea usually moderately deep and teeth moderately robust [fig.19]; median lobe of aedeagus with auricles smaller, not uncinate to moderately uncinate; body small to medium sized (length 1.6-2.8 mm); scales of elytral interstriae often arranged in single regular rows; uncus of protibia in male small.

   ...16

MIARUS URSINUS
fig.19
 
#16 Pronotum moderately convex on disc; median lobe of aedeagus with auricles not uncinate and with tongue distinctly shorter [fig.40]. South-western Europe: Portugal, Spain, France (incl. Corsica), Switzerland, peninsular Italy, Croatia..

   ...ursinus Abeille, 1906

MIARUS URSINUS.GIF
fig.40
 
-- Pronotum usually distinctly convex on disc; median lobe of aedeagus with auricles uncinate and with tongue distinctly longer [fig.46]. North-western Africa: Morocco, Algeria..

   ...maroccanus Solari, 1947

MIARUS MAROCCANUS.GIF
fig.46
 
#17 Median lobe of aedeagus with distinct mitre recurved posteriorly [fig.47]. Russia (Far East), Kazakhstan, Kyrgyzstan, China, Japan, Korea.

   ...atricolor Morimoto, 1983

MIARUS ATRICOLOR.GIF
fig.47
 
-- Median lobe of aedeagus without or with very small mitre.

   ...18

 
#18 Median lobe of aedeagus with mitre and auricles small but distinct, with anteapical stricture very strong, with tongue larger in basal half and clearly visible in dorsal view [fig.43]. France, Switzerland, Poland, Germany, Czech Republic, Slovakia, Hungary, Austria, Italy, Slovenia, Croatia, Bosnia and Herzegovina, Serbia, Montenegro, Macedonia, Armenia, Denmark, Sweden, Norway, Finland, Russia (from North West to Far East), Mongolia, Korea, Syria, Iran, north-western Africa.

   ...ajugae (Hbst., 1795)

MIARUS AJUGAE3.GIF
fig.43
 
-- Median lobe of aedeagus without mitre and auricles, with anteapical stricture moderately strong, with tongue smaller in basal half and indistinct in dorsal view [fig.42]. Spain, France, northern and central Italy, Serbia, Romania, Bulgaria, European Turkey, Georgia.

Meldungen in Google Earth anzeigen   ...simplex Solari, 1947

MIARUS SIMPLEX.GIF
fig.42
 
  abeillei
abnormis
afer
ajugae
araxis
atricolor
banaticus
campanulae
dentiventris
hellenicus
Iongicollis
maroccanus
monticola
praecursor
rotundicollis
simplex
solarii
stoeckleini
ursinus
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First prepared: 28.12.2015
Latest edit: 31.03.2019 - 00:59:51
Käfer Europas by Arved Lompe / Miarus by Roberto Caldara
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