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Gattung Ernobius |
Coleoptera - Teredilia - Anobiidae |
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By Colin Johnson (Rearranged by: Arved Lompe) If you find any errors or omissions please mail me at mailbox@lompe.de |
Johnson, C. (1975): A Review of the palaearctic Species of the Genus Ernobius Thomson - Ent.Bl. 71(2):65-93 Private Datei: F:\taxa\coleo\scans\Johnson1975 Ernobius.pdf |
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Species not listed in this key ...99 |
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#1 |
Antennal segments 6-8 very short and slender, together upto half the length of segment 9, and shorter than 3-5 combined (sometimes almost as long in laticollis ♀). ...2 |
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Antennal segments 6-8 never so short, frequently quite thick, together always longer than segment 9, and as long or longer than 3-5 combined. ...13 |
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#2 |
Very small and narrow brown species, 1.8-2.8 mm. ...3 |
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Larger and broader species, over 3 mm in length. ...4 |
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#3 |
Front angles of the pronotum distinct, due to the sides being strongly sinuate (as in longicornis); pronotum not explanate at the sides. Aedeagus [fig.1]. ...juniperi Chob. |
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Front angles of the pronotum effaced, the sides close to them not sinuate; pronotum explanate at the sides. Aedeagus [fig.2]. ...cupressi Chob. |
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#4 |
Larger species, 4.5-7.0 mm; surface rather dull; pronotum and elytra coarsely and closely granulate, the granules weakly punctate, the sculpture similar on both; elytra strongly impressed near the shoulders and on each side of the scutellum; dark reddish-brown species. ...5 |
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Smaller species, 3.0-4.0 mm; surface more shining; pronotum with less coarse and conspicuously punctate granules; elytra more finely and sparsely granulate or granulo-punctate, the sculpture dissimilar on both; elytra not or feebly impressed; colour typically entirely black, the elytra often lighter, ocasionally entirely dark brown. ...6 |
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#5 |
Aedeagus [fig.3]. ...laticollis Pic |
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Aedeagus [fig.4]. ...reitteri Pic |
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#6 |
Base of the pronotum with a transverse depression. ...7 |
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Base of the pronotum without a transverse depression. ...9 |
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#7 |
Front angles of the pronotum distinct, due to the sides before them being strongly sinuate [fig.5] [fig.6]; elytra more elongate, c. 2.5 times as long as broad [fig.7]; antennae reddish-brown; pronotum as broad (♀) or narrower (♂) than the elytra. Aedeagus [fig.8] [fig.9]. ...longicornis (Sturm, 1837) |
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Front angles of the pronotum effaced, the sides not sinuate; elytra not so elongate, c. twice as long as broad; antennae usually black. ...8 |
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#8 |
Pronotum narrower than the elytra; eyes strongly protruding. Aedeagus [fig.10]. ...freudei Lohse |
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Pronotum as broad as the elytra; eyes not quite so strongly protruding. Habitus [fig.11]. Aedeagus [fig.12]. ...nigrinus (Sturm, 1837) |
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#9 |
Entirely black, except for the elytral apex which is bright reddish-yellow; antennae black, segment 10 obvious bent; segments 6-8 extremely short in ♂. Aedeagus [fig.13]. ...pardoi Espanol |
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Elytra always unicolorous, various shades of brown; antennae usually brown, very rarely black, segment 10 not or very feebly bent, 6-8 not so short. ...10 |
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#10 |
Base of the pronotum clearly sinuate on each side of the middle, the curvature thus uneven; pronotum sparsely covered with larger umbilicate punctures which are predominantly C-shaped, the surface shining. ...11 |
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Base of the pronotum not or barely sinuate on each side of the middle, the base and sides thus evenly and continuously curved; pronotum very closely covered with finer umbilicate punctures which are almost circular, the surface somewhat dull; brown species. ...12 |
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#11 |
Bright yellowish-brown, antennae included; sides of the pronotum less curved apically, narrowly explanate. Aedeagus [fig.14]. ...kiesenwetteri Schilsky, 1899 |
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Brown to brown-black, antennae included; sides of the pronotum more curved apically, narrowly explanate. Aedeagus [fig.15]. ...pallidipennis Pic |
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#12 |
Size larger, 4 mm. Aedeagus [fig.16]. ...subopacus Pic Size smaller, 3.0-3.5 mm. Aedeagus [fig.17]. ...syriacus Pic |
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#13 |
Dorsal surface of tarsal segment 4 with the apical excavation not or barely surpassing the middle [fig.18]; mostly larger species, 4.0-6.0 mm, very rarely as small as 3.0 mm. Tarsal segment 5 long and narrow in most species. Aedeagus of a similar basic form [fig.19]. ...14 |
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Dorsal surface of tarsal segment 4 with the apical excavation clearly surpassing the middle, generally occupying at least three-quarters of the length of the segment [fig.20] [fig.21]; mostly smaller species, 2.0- 4.0 mm, very rarely upto 5.0 mm. Aedeagi of various different forms. ...19 |
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#14 |
Front tibiae conspicuously curved inwards (more so in ♂), at least in their basal half, the apex not or barely turned outwards. ...15 Front tibiae straight, the apex clearly turned outwards. ...16 |
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#15 |
Pronotum broadly expIanate at the sides; antennae rather short and thick, segments 6-8 from one to 1.5 times as long as broad; body rather slender; colour usually very dark brown; average size 4.0 mm; tarsal segment 5 long in ♂, short in ♀. Aedeagus [fig.19]. North Europe. ...explanatus (Mannh.) |
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Hooked paramere without a median bulge [fig.22]. Central Europe ssp. ...phobos Gottw. |
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Pronotum narrowly explanate at the sides; antennae long and slender, segments 6-8 at least twice as long as broad; body rather broad; colour yellowish / reddish-brown; average size 5.0 mm; tarsal segment 5 long in both sexes. Aedeagus [fig.23]. ...gigas Mulsant & Rey, 1863 |
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#16 |
Bright yellowish-brown, very shining; pronotum sparsely covered with umbilicate punctures which are C-shaped to nearly circular; pronotal disc longitudinally convex; antennae similar to reflexus. Aedeagus [fig.24]. ...lucidus (Muls. & Rey) |
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Reddish-brown species, not so brilliantly shining; pronotum very closely covered with finer punctures or granules, umbilicate or otherwise, which are predominantly circular; pronotal disc longitudinally flatter. ...17 |
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#17 |
Size larger, 5.0-6.0 mm; pronotum very broadly explanate at the sides, which are strongly rounded like the base; pronotal punctures conspicuously umbilicate, more-or-less circular; antenna! segment 8 over half (usually two thirds) the length of 9; pubescence longer, more golden and less depressed than in the next species. Aedeagus [fig.25]. ...reflexus (Muls. & Rey) |
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Size smaller, 3.0-5.0 mm; pronotum not or only narrowly explanate at the sides, which are not as strongly rounded; pronotal sculpture more granular, not or weakly umbilicate; antenna! segment 8 under half the length of 9 . ...18 |
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#18 |
Pronotum finely granulate, not umbilicate [fig.26]. Aedeagus [fig.27] [fig.28]. ...mollis (L., 1758) |
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Pronotum coarsely granulate, the granules weakly but distinctly umbilicate. Pointed paramere [fig.29]. Tenerife . mollis ssp. ...espanoli Johnson, 1975 |
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#19 |
Posterior margin of pronotum strongly oblique on either side, hind angles some distance from elytral base [fig.30]; pronotum narrower than elytra, front angles sinuate and distinct, dorsum black except for the brown elytra (♂), pronotum as broad as elytra, front angles effaced and not sinuate, dorsum unicolorous brown (♀); tarsal segment 5 short and broad. Aedeagus [fig.31]. ...angusticollis (Ratz., 1847) [fig.32] |
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Posterior margin of pronotum more-or-less straight, never so strongly oblique on each side, the hind angles more-or-less touching the elytral base; most species entirely brown in both sexes. ...20 |
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#20 |
Pronotum very dull, with strong reticulation, very closely covered with fine granules which are circular and barely umbilicate. Antenna! segment 8 generally over three-quarters (very rarely barely over half) the length of 9; body long and narrow; pronotum narrower than the elytra (♂), or as broad (♀). Aedeagus [fig.33]. ...pruinosus (Muls. Rey) |
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Pronotum more shining, without or rarely with weaker reticulation, covered with somewhat larger circular or C-shaped umbilicate punctures / granules. ...21 |
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#21 |
Antenna! segments 7 and 8 somewhat similar to 9-11, ie. segment 8 is usually at least three-quarters the length of 9; body long and narrow in one species; pronotum narrower than elytra (♂) or as broad (♀). ...22 |
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Antenna! segments 7 and 8 different from 9-11 , ie. segment 8. barely exceeds half the length of 9; body always shorter. ...23 |
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#22 |
Size smaller, 2.0-3.0 mm; body long and narrow; pronotum somewhat sparsely umbilicate-granulate, these almost circular; antennae and tarsi more slender. Aedeagus [fig.34] [fig.35]. ...abietinus (Gyll., 1808) |
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Size larger, 3.3-4.0 mm; body shorter and broader; pronotum densely umbilicate-granulate, these circular; antennae and tarsi thicker and broader. Aedeagus [fig.36]. ...oxycedri Reitt. |
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#23 |
Tarsal segment 5 rather short and broad, the protruding part barely exceeding 1.5 times its greatest breadth in length [fig.20]; tarsi more-or- less broad. ...24 |
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Tarsal segment 5 long and slender, the protruding part 2-3 times its greatest breadth in length [fig.21]; tarsi narrow. ...30 |
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#24 |
Tarsal segment 5 very short, the protruding part about as long as its greatest breadth; antenna! segment 9 as long as segments 6-8 combined. Habitus [fig.37], detail of front body [fig.38]. Aedeagus [fig.39]. ...abietis (F., 1792) |
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Tarsal segment 5 not so short, the protruding part almost 1.5 times as long as its greatest breadth; antenna! segment 9 shorter in length than 6-8 combined, generally only as long as 7-8. ...25 |
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#25 |
East Palaearctic species (Japan); antennae short, rather slender in both sexes. Aedeagus [fig.40]. ...curticollis Pic |
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West Palaearctic species; antennae longer and thicker, often considerably so in the male. ...26 |
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#26 |
Elytra very coarsely and densely granulate. ...27 |
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Elytra finely and sparsely granulo-punctate. ...28 |
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#27 |
Elytral pubescence rather long and semi-erect; pronotum shining, closely covered with umbilicate granules which tend to be more C-shaped than circular, except towards the front; antennal segment 6 shorter than 7. Aedeagus [fig.41]. ...robusticornis Maran |
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Elytral pubescence shorter and much more depressed as in most species; pronotum rather dull, very closely covered with smaller, umbilicate and circular granules which are very closely packed together; antenna! segment 6 about as long as 7. Aedeagus [fig.42]. ...kaiIidisi Johnson, 1975 |
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#28 |
Elytra unicolorous dark brown, its pubescence more-or-less directed rearwards; head almost as broad as the pronotum, the pronotum narrower than the elytra in the ♂. Aedeagus [fig.43]. ...29 |
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Elytra brown with a black stripe down the suture; elytral pubescence directed inwards from the sides and sutural area, more-or-less rearwards between; head narrower than the pronotum which is as broad as the elytra in the ♂. Aedeagus [fig.44]. ...anatoIicus Johnson, 1975 |
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#29 |
Size smaller, 3.0-4.0 mm; antenna! segments shorter [fig.45]; pronotum closely covered with umbilicate granules. Aedeagus [fig.43]. Europe. ...mulsanti Kiesw., 1877 |
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Size larger, 4.0-5.5 mm; antenna! segments more elongate [fig.46]; pronotum very densely packed with umbilicate granules. North Africa mulsanti ssp. ...fructuum Peyerh. |
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#30 |
Major part of the pronotum conspicuously punctured; towards the front there are indistinct C-shaped granules around the punctures; body long and narrow; pronotum narrower than the elytra (♀); dark brown. Cyprus. ...madoni Pic |
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Pronotum clearly granulate, the granules generally obviously umbilicate; body shorter and broader; chiefly paler brown species. ...31 |
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#31 |
Umbilicate (-punctate) granules on the centre of the pronotum predominantly C-shaped. ...32 |
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Umbilicate granules on the centre of the pronotum predominantly circular. ...33 |
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#32 |
Bright yellowish-brown, very shining as a rule; pronotum broadly explanate at the sides, the hind angles strongly rounded off; elytral pubescence somewhat long and semi-erect; antennae rather long and thick. Aedeagus [fig.47]. ...rufus (Illig.) |
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Reddish-brown, somewhat dull, pronotum narrowly explanate at the sides, the hind angles more angularly curved [fig.48]; elytral pubescence shorter and more depressed; antennae shorter and more slender. Aedeagus [fig.49] [fig.50]. ...pini (Sturm, 1837) |
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#33 |
Pronotum reticulate, broadly explanate at the sides, including the front angles which are somewhat prominent; elytra dull, finely and very closely granulate; antenna! segment 9 longer than 7 and 8 together, but shorter than 6-8 together, Aedeagus [fig.51]. ...impressithorax Pic |
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Pronotum at the most narrowly explanate at the sides, the explanation having petered out at the front angles, which are effaced; elytra more shining, not quite so closely granulate; antenna} segment 9 only as long as segments 7 and 8 together. ...34 |
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#34 |
Average ,size larger, 3.0-3.5 mm, very rarely down to 2.5 mm; pronotum generally somewhat dull due to being finely but clearly reticulate; pronotum closely covered with fine umbilicate granules. Aedeagus [fig.52]. ...oertzeni Schils. |
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Average size smaller, 1.7-3.0 mm; pronotum usually rather shining, only very rarely reticulate. ...35 |
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#35 |
Sides of the pronotum clearly explanate and thus somewhat flattened almost upto the front angles; pronotum shorter, generally with larger and more scattered umbilicate granules. Aedeagus [fig.53]. ...parens Muls. & Rey |
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Sides of the pronotum hardly explanate at any rate in front of the middle, not or barely flattened except near the hind angles; pronotum generally a trifle longer, with slightly smaller and more dense umbilicate granules. ...36 |
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#36 |
Antennae [fig.54] slightly thicker; pronotum conspicuously umbilicate; dorsum black, except for the brown elytra (♂), dorsum brown (♀). Aedeagus [fig.55]. ...gallicus Johnson, 1975 |
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Antennae [fig.56] more slender; pronotum weakly umbilicate; dorsum entirely brown in both sexes. Aedeagus [fig.57]. ...fulvus Johnson, 1975 |
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#99 |
France. ...oculeus Toskina, 2002 |
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Spain. ...vinolasi Novoa & Baselga, 2000 |
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Switzerland. ...besucheti Zahradnik, 2000 |
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Italy. ...angelinii Lohse, 1991 |
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Russia. ...jaroslavli Logvinovskiy, 1977 |
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abietinus abietis angelinii angusticollis besucheti cupressi curticollis espanoli explanatus freudei fructuum fulvus gallicus gigas impressithorax jaroslavli juniperi kiesenwetteri laticollis longicornis lucidus madoni |
mollis mulsanti nigrinus oculeus oertzeni oxycedri pallidipennis pardoi parens phobos pini pruinosus reflexus reitteri robusticornis rufus subopacus syriacus vinolasi anatoIicus kaiIidisi |
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First prepared: Arved Lompe) > |
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